About the Authors

Stephen N. Sansom

Affiliation Gurdon Institute and Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Dean S. Griffiths

Contributed equally to this work with: Dean S. Griffiths, Andrea Faedo

Current address: Cambridge Institute for Medical Research, Cambridge, United Kingdom

Affiliation Gurdon Institute and Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Andrea Faedo

Contributed equally to this work with: Dean S. Griffiths, Andrea Faedo

Affiliation Nina Ireland Laboratory of Developmental Neurobiology, Department of Psychiatry, University of California at San Francisco, San Francisco, California, United States of America

Dirk-Jan Kleinjan

Affiliation Medical Research Council Human Genetics Unit, Western General Hospital, Edinburgh, United Kingdom

Youlin Ruan

Affiliation Nina Ireland Laboratory of Developmental Neurobiology, Department of Psychiatry, University of California at San Francisco, San Francisco, California, United States of America

James Smith

Affiliation Gurdon Institute and Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Veronica van Heyningen

Affiliation Medical Research Council Human Genetics Unit, Western General Hospital, Edinburgh, United Kingdom

John L. Rubenstein

Affiliation Nina Ireland Laboratory of Developmental Neurobiology, Department of Psychiatry, University of California at San Francisco, San Francisco, California, United States of America

Frederick J. Livesey

rick@gurdon.cam.ac.uk

Affiliation Gurdon Institute and Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SNS FJL. Performed the experiments: SNS DSG AF JS. Analyzed the data: SNS DSG AF JLR FJL. Contributed reagents/materials/analysis tools: DJK YR VvH JLR. Wrote the paper: SNS FJL.