About the Authors

Charles T. Miller

Affiliations Graduate Program in Cell and Molecular Biology, Michigan State University, East Lansing, Michigan, United States of America, Laboratory of Chromosome Replication, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Carrie Gabrielse

Affiliation Laboratory of Chromosome Replication, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Ying-Chou Chen

Affiliations Laboratory of Chromosome Replication, Van Andel Research Institute, Grand Rapids, Michigan, United States of America, Graduate Program in Genetics, Michigan State University, East Lansing, Michigan, United States of America

Michael Weinreich

michael.weinreich@vai.org

Affiliation Laboratory of Chromosome Replication, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MW. Performed the experiments: CTM CG YCC MW. Analyzed the data: CTM CG YCC MW. Contributed reagents/materials/analysis tools: CTM CG YCC MW. Wrote the paper: MW.