About the Authors

Michael A. Nalls

Affiliations Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America, Laboratory of Epidemiology, Demography and Biometry, Intramural Research Program, National Institute on Aging, Maryland, United States of America

Javier Simon-Sanchez

Affiliations Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America, Unidad de Genética Molecular, Departamento de Genómica y Proteómica, Instituto de Biomedicina de Valencia-CSIC, Valencia, Spain

J. Raphael Gibbs

Affiliations Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America, Department of Molecular Neuroscience and Reta Lila Weston Laboratories, Institute of Neurology, London, United Kingdom

Coro Paisan-Ruiz

Affiliation Department of Molecular Neuroscience and Reta Lila Weston Laboratories, Institute of Neurology, London, United Kingdom

Jose Tomas Bras

Affiliation Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America

Toshiko Tanaka

Affiliation Longitudinal Studies Section, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America

Mar Matarin

Affiliation Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America

Sonja Scholz

Affiliations Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America, Department of Molecular Neuroscience and Reta Lila Weston Laboratories, Institute of Neurology, London, United Kingdom

Charles Weitz

Affiliation Department of Anthropology, Biological Anthropology Program, Temple University, Philadelphia, Pennsylvania, United States of America

Tamara B. Harris

Affiliation Laboratory of Epidemiology, Demography and Biometry, Intramural Research Program, National Institute on Aging, Maryland, United States of America

Luigi Ferrucci

Affiliation Longitudinal Studies Section, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America

John Hardy

Affiliation Department of Molecular Neuroscience and Reta Lila Weston Laboratories, Institute of Neurology, London, United Kingdom

Andrew B. Singleton

singleta@mail.nih.gov

Affiliations Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America, Department of Public Health Sciences, Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JH ABS. Performed the experiments: JSS CPR SS. Analyzed the data: MAN JSS JRG TT CW LF ABS. Contributed reagents/materials/analysis tools: JSS JTB TT MM CW TBH LF JH ABS. Wrote the paper: MAN JSS ABS.