About the Authors

Patrícia Beldade

p.beldade@biology.leidenuniv.nl

Affiliations Institute of Biology, Leiden University, Leiden, The Netherlands, Instituto Gulbenkian de Ciência, Oeiras, Portugal

Suzanne V. Saenko

Affiliation Institute of Biology, Leiden University, Leiden, The Netherlands

Nicolien Pul

Affiliation Institute of Biology, Leiden University, Leiden, The Netherlands

Anthony D. Long

Affiliation Department of Ecology and Evolutionary Biology, University of California Irvine, Irvine, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PB ADL. Performed the experiments: PB ADL. Analyzed the data: PB SVS ADL. Contributed reagents/materials/analysis tools: PB SVS NP ADL. Wrote the paper: PB ADL. Provided the material for and assisted in mapping the visible markers: SVS. Prepared the butterfly material and identified the SNPs in candidate genes: NP.