About the Authors

Sergey Kryazhimskiy

Affiliation Biology Department, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Joshua B. Plotkin

jplotkin@sas.upenn.edu

Affiliations Biology Department, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Program in Applied Mathematics and Computational Science, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JBP. Performed the experiments: SK. Analyzed the data: SK. Wrote the paper: SK JBP. Designed the research: SK JBP.