About the Authors

Alkes L. Price

aprice@hsph.harvard.edu (ALP); spielman@pobox.upenn.edu (RSS)

Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America

Nick Patterson

Affiliation Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Dustin C. Hancks

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Simon Myers

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

David Reich

Affiliations Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Vivian G. Cheung

Affiliations Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Pediatrics, University of Pennsylvania School of Medicine, Pennsylvania, United States of America, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America, Howard Hughes Medical Institute, Philadelphia, Pennsylvania, United States of America

Richard S. Spielman

aprice@hsph.harvard.edu (ALP); spielman@pobox.upenn.edu (RSS)

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ALP NP DR VGC RSS. Performed the experiments: DCH VGC RSS. Analyzed the data: ALP NP SM DR VGC RSS. Wrote the paper: ALP DR VGC RSS. Contributed data: DCH VGC RSS.