About the Authors

Sang Hong Lee

Affiliations School of Environmental and Rural Science, University of New England, Armidale, New South Wales, Australia, National Institute of Animal Science, Rural Development Administration, Cheon An, Korea

Julius H. J. van der Werf

Affiliation School of Environmental and Rural Science, University of New England, Armidale, New South Wales, Australia

Ben J. Hayes

Affiliation Department of Primary Industry, Victoria, Australia

Michael E. Goddard

Affiliations Department of Primary Industry, Victoria, Australia, Faculty of Land and Food Resources, University of Melbourne, Melbourne, Australia

Peter M. Visscher

peter.visscher@qimr.edu.au

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SHL JHJvdW MEG PMV. Analyzed the data: SHL. Contributed reagents/materials/analysis tools: JHJvdW BJH. Wrote the paper: SHL JHJvdW MEG PMV.