About the Authors

Michael F. Hammer

mfh@u.arizona.edu

Affiliations ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America, Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Arizona, United States of America

Fernando L. Mendez

Affiliation Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Arizona, United States of America

Murray P. Cox

Affiliation ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America

August E. Woerner

Affiliation ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America

Jeffrey D. Wall

Affiliation Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MFH JDW. Performed the experiments: MPC AEW. Analyzed the data: MFH FLM MPC AEW JDW. Wrote the paper: MFH FLM JDW.