About the Authors

Yoshito Oka

Contributed equally to this work with: Yoshito Oka, Tomonao Matsushita

Affiliations Department of Biology, Graduate School of Science, Kyoto University, Kyoto, Japan, Department of Plant and Microbial Biology, University of California Berkeley, Berkeley, California, United States of America, Plant Gene Expression Center, U.S. Department of Agriculture, Albany, California, United States of America

Tomonao Matsushita

Contributed equally to this work with: Yoshito Oka, Tomonao Matsushita

Affiliations Department of Biology, Graduate School of Science, Kyoto University, Kyoto, Japan, Graduate School of Agriculture, Kyushu University, Fukuoka, Japan, Organization for the Promotion of Advanced Research, Kyushu University, Fukuoka, Japan

Nobuyoshi Mochizuki

Affiliation Department of Biology, Graduate School of Science, Kyoto University, Kyoto, Japan

Peter H. Quail

Affiliations Department of Plant and Microbial Biology, University of California Berkeley, Berkeley, California, United States of America, Plant Gene Expression Center, U.S. Department of Agriculture, Albany, California, United States of America

Akira Nagatani

nagatani@physiol.bot.kyoto-u.ac.jp

Affiliation Department of Biology, Graduate School of Science, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YO TM PHQ AN. Performed the experiments: YO TM NM AN. Analyzed the data: YO TM PHQ AN. Contributed reagents/materials/analysis tools: NM PHQ. Wrote the paper: YO TM PHQ AN.