About the Authors

Monica Chang

Affiliation Celera, Alameda, California, United States of America

Charles M. Rowland

Affiliation Celera, Alameda, California, United States of America

Veronica E. Garcia

Affiliation Celera, Alameda, California, United States of America

Steven J. Schrodi

Affiliation Celera, Alameda, California, United States of America

Joseph J. Catanese

Affiliation Celera, Alameda, California, United States of America

Annette H. M. van der Helm-van Mil

Affiliation Leiden University Medical Centre, Leiden, The Netherlands

Kristin G. Ardlie

Address Current address: Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Affiliation SeraCare Life Sciences, Cambridge, Massachusetts, United States of America

Christopher I. Amos

Affiliation University of Texas, Houston, Texas, United States of America

Lindsey A. Criswell

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Daniel L. Kastner

Affiliation National Institute of Health, Bethesda, Maryland, United States of America

Peter K. Gregersen

Affiliation Feinstein Institute for Medical Research, North Shore L.I.J. Health System, Manhasset, New York, United States of America

Fina A. S. Kurreeman

Affiliation Leiden University Medical Centre, Leiden, The Netherlands

Rene E. M. Toes

Affiliation Leiden University Medical Centre, Leiden, The Netherlands

Tom W. J. Huizinga

Affiliation Leiden University Medical Centre, Leiden, The Netherlands

Michael F. Seldin

Affiliation University of California Davis, Davis, California, United States of America

Ann B. Begovich

ann.begovich@roche.com

Address Current address: Roche Diagnostics, Pleasanton, California, United States of America

Affiliation Celera, Alameda, California, United States of America

Competing Interests

MC, CMR, VEG, SJS, JJC and ABB are employees of Celera and, as such, may own stock and/or stock options in Celera.

Author Contributions

Conceived and designed the experiments: SS AB. Performed the experiments: MC JC. Analyzed the data: MC CR VG SS AB. Contributed reagents/materials/analysis tools: CR VG SS AvdH KA CA LC DK PG FK RT TH MS. Wrote the paper: CR SS AB.