About the Authors

Hongkui Zeng

Contributed equally to this work with: Hongkui Zeng, Kyoji Horie

Address Current address: Allen Institute for Brain Science, Seattle, Washington, United States of America;

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Kyoji Horie

Contributed equally to this work with: Hongkui Zeng, Kyoji Horie

Address Current address: Department of Social and Environmental Medicine, Graduate School of Medicine, Osaka University, Osaka, Japan.

Affiliation Human Retrovirus Section, Vaccine Branch, Center for Cancer Research, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

Linda Madisen

Address Current address: Allen Institute for Brain Science, Seattle, Washington, United States of America;

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Maria N. Pavlova

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Galina Gragerova

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Alex D. Rohde

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Brian A. Schimpf

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Yuqiong Liang

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Ethan Ojala

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Farah Kramer

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Patricia Roth

Affiliation Human Retrovirus Section, Vaccine Branch, Center for Cancer Research, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

Olga Slobodskaya

Affiliation Human Retrovirus Section, Vaccine Branch, Center for Cancer Research, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

Io Dolka

Affiliation Human Retrovirus Section, Vaccine Branch, Center for Cancer Research, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

Eileen A. Southon

Affiliation Neural Development Section, Mouse Cancer Genetics Program, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

Lino Tessarollo

Affiliation Neural Development Section, Mouse Cancer Genetics Program, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

Karin E. Bornfeldt

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Alexander Gragerov

Affiliation Omeros Corporation, Seattle, Washington, United States of America

George N. Pavlakis

ggaitanaris@omeros.com (GGA); pavlakis@ncifcrf.gov (GNP)

Affiliation Human Retrovirus Section, Vaccine Branch, Center for Cancer Research, National Cancer Institute-Frederick, Frederick, Maryland, United States of America

George A. Gaitanaris

ggaitanaris@omeros.com (GGA); pavlakis@ncifcrf.gov (GNP)

Affiliation Omeros Corporation, Seattle, Washington, United States of America

Competing Interests

Authors currently affiliated with Omeros Corporation wish to declare potential financial interests.

Author Contributions

Conceived and designed the experiments: HZ KH LM LT KB GP GAG. Performed the experiments: HZ KH LM MP GG AR BS YL EO FK PR OS ID ES AG. Analyzed the data: HZ KH KB AG GAG. Contributed reagents/materials/analysis tools: FK ES LT GP. Wrote the paper: HZ KH GAG.