About the Authors

Kazuma Hanai

Affiliations Department of Developmental Genetics, National Institute of Genetics, Shizuoka-ken, Japan, Department of Mathematical and Life Science, Graduate School of Science, Hiroshima University, Higashi-Hiroshima, Japan

Hirofumi Furuhashi

Address Current address: Department of Biology, O. Wayne Rollins Research Center, Emory University, Atlanta, Georgia, United States of America

Affiliation Department of Developmental Genetics, National Institute of Genetics, Shizuoka-ken, Japan

Takashi Yamamoto

Affiliation Department of Mathematical and Life Science, Graduate School of Science, Hiroshima University, Higashi-Hiroshima, Japan

Koji Akasaka

Affiliation Misaki Marine Biological Station, Graduate School of Science, The University of Tokyo, Miura, Kanagawa, Japan

Susumu Hirose

shirose@lab.nig.ac.jp

Affiliation Department of Developmental Genetics, National Institute of Genetics, Shizuoka-ken, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HF TY KA SH. Performed the experiments: KH HF. Analyzed the data: KH HF SH. Contributed reagents/materials/analysis tools: KH. Wrote the paper: KH SH.