About the Authors

Changqi C. Zhu

Contributed equally to this work with: Changqi C. Zhu, Douglas J. Bornemann

Affiliations Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Douglas J. Bornemann

Contributed equally to this work with: Changqi C. Zhu, Douglas J. Bornemann

Affiliations Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

David Zhitomirsky

Affiliations Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Ellen L. Miller

Affiliation Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Michael B. O'Connor

Affiliations Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Jeffrey A. Simon

simon004@umn.edu

Affiliation Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CZ DB MO JS. Performed the experiments: CZ DB DZ EM MO JS. Analyzed the data: CZ DB MO JS. Contributed reagents/materials/analysis tools: CZ DB MO JS. Wrote the paper: CZ DB MO JS.