About the Authors

William G. Hill
Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Michael E. Goddard
Faculty of Land and Food Resources, University of Melbourne, Victoria, Australia

Michael E. Goddard
Department of Primary Industries, Victoria, Australia

Peter M. Visscher
Queensland Institute of Medical Research, Brisbane, Australia

Corresponding Author

w.g.hill@ed.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WH MG PV. Analyzed the data: WH PV. Wrote the paper: WH MG PV.