About the Authors

William G. Hill


Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Michael E. Goddard

Affiliations Faculty of Land and Food Resources, University of Melbourne, Victoria, Australia, Department of Primary Industries, Victoria, Australia

Peter M. Visscher

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WH MG PV. Analyzed the data: WH PV. Wrote the paper: WH MG PV.