About the Authors

Jonathan S Friedlaender

To whom correspondence should be addressed. E-mail: jfriedla@temple.edu

Affiliation Anthropology Department, Temple University, Philadelphia, Pennsylvania, United States of America

Françoise R Friedlaender

Affiliation Independent Researcher, Philadelphia, Pennsylvania, United States of America

Floyd A Reed

Affiliation Department of Biology, University of Maryland, College Park, Maryland, United States of America

Kenneth K Kidd

Affiliation Department of Genetics, Yale University, New Haven, Connecticut, United States of America

Judith R Kidd

Affiliation Department of Genetics, Yale University, New Haven, Connecticut, United States of America

Geoffrey K Chambers

Affiliation School of Biological Sciences, Victoria University, Wellington, New Zealand

Rodney A Lea

Affiliation School of Biological Sciences, Victoria University, Wellington, New Zealand

Jun-Hun Loo

Affiliation Transfusion Medicine Laboratory, Mackay Memorial Hospital, Taipei, Taiwan

George Koki

Affiliation Institute for Medical Research, Goroka, Eastern Highlands Province, Papua New Guinea

Jason A Hodgson

Address ¤ Current address: Center for the Study of Human Origins and the New York Consortium in Evolutionary Primatology, Department of Anthropology, New York University, New York, New York, United States of America

Affiliation Department of Anthropology, Binghamton University, Binghamton, New York, United States of America

D. Andrew Merriwether

Affiliation Department of Anthropology, Binghamton University, Binghamton, New York, United States of America

James L Weber

Affiliation Marshfield Clinic Research Foundation, Marshfield, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JSF and DAM designed the study. JSF, GK, JRK, KKK, GKC, RAL, J-HL, and DAM collected or contributed samples, which were prepared for genotyping by JAH. JLW supervised genotyping at the Marshfield Clinic. The data analysis was performed by FRF, with advice and assistance from FAR, JSF, JRK, and KKK. JSF and FRF wrote the text.