About the Authors

Jonathan S Friedlaender
Anthropology Department, Temple University, Philadelphia, Pennsylvania, United States of America

Françoise R Friedlaender
Independent Researcher, Philadelphia, Pennsylvania, United States of America

Floyd A Reed
Department of Biology, University of Maryland, College Park, Maryland, United States of America

Kenneth K Kidd, Judith R Kidd
Department of Genetics, Yale University, New Haven, Connecticut, United States of America

Geoffrey K Chambers, Rodney A Lea
School of Biological Sciences, Victoria University, Wellington, New Zealand

Jun-Hun Loo
Transfusion Medicine Laboratory, Mackay Memorial Hospital, Taipei, Taiwan

George Koki
Institute for Medical Research, Goroka, Eastern Highlands Province, Papua New Guinea

Jason A Hodgson, D. Andrew Merriwether
Department of Anthropology, Binghamton University, Binghamton, New York, United States of America

James L Weber
Marshfield Clinic Research Foundation, Marshfield, Wisconsin, United States of America

Corresponding Author


Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JSF and DAM designed the study. JSF, GK, JRK, KKK, GKC, RAL, J-HL, and DAM collected or contributed samples, which were prepared for genotyping by JAH. JLW supervised genotyping at the Marshfield Clinic. The data analysis was performed by FRF, with advice and assistance from FAR, JSF, JRK, and KKK. JSF and FRF wrote the text.