About the Authors

Alkes L Price, David Reich, Joel N Hirschhorn
Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Alkes L Price, Johannah Butler, Nick Patterson, Alicja Waliszewska, Christine Schirmer, Kristin Ardlie, Alexis Ramos, James Nemesh, Lori Arbeitman, David Reich, Joel N Hirschhorn
Program in Medical and Population Genetics, Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Johannah Butler, Alexis Ramos, James Nemesh, Lori Arbeitman, Joel N Hirschhorn
Program in Genomics and Divisions of Genetics and Endocrinology, Children's Hospital, Boston, Massachusetts, United States of America

Cristian Capelli
Department of Zoology, University of Oxford, Oxford, United Kingdom

Vincenzo L Pascali, Francesca Scarnicci
Forensic Genetics Laboratory, Istituto di Medicina Legale, Universita Cattolica del Sacro Cuore, Rome, Italy

Andres Ruiz-Linares
Department of Biology, Galton Laboratory, University College London, United Kingdom

Leif Groop
Department of Clinical Sciences, Diabetes and Endocrinology, Lund University, University Hospital Malmo, Malmo, Sweden

Angelica A Saetta, Penelope Korkolopoulou
Department of Pathology, Medical School, National and Kapodistrian University of Athens, Athens, Greece

Uri Seligsohn
Amalia Biron Research Institute of Thrombosis and Hemostasis, Sheba Medical Center, Tel Hashomer, Israel

David B Goldstein
Institute for Genome Sciences and Policy, Center for Population Genomics and Pharmacogenetics, Duke University, Durham, North Carolina, United States of America

Corresponding Authors

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

ALP, JB, DR, and JNH conceived and designed the experiments. ALP, JB, AW, CS, AR, JN, and LA performed the experiments. ALP analyzed the data. ALP, NP, CC, VLP, FS, ARL, LG, AAS, PK, US, KA, DBG, DR, and JNH contributed reagents/materials/analysis tools. ALP, DR, and JNH wrote the paper.