About the Authors

Yan Yang

Contributed equally to this work with: Yan Yang, Connie L Mahaffey

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Connie L Mahaffey

Contributed equally to this work with: Yan Yang, Connie L Mahaffey

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Nathalie Bérubé

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Terry P Maddatu

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Gregory A Cox

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Wayne N Frankel

To whom correspondence should be addressed. E-mail: wayne.frankel@jax.org

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

YY and WNF conceived and designed the experiments and wrote the paper. YY, CLM, NB, and WNF performed the experiments and analyzed the data. TPM and GAC contributed reagents/materials/analysis tools.