About the Authors

Araceli G Castillo

Contributed equally to this work with: Araceli G Castillo, Barbara G Mellone

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Barbara G Mellone

Contributed equally to this work with: Araceli G Castillo, Barbara G Mellone

Address Current address: Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Janet F Partridge

Address Current address: Department of Biochemistry, St. Jude Children's Research Hospital, Memphis, Tennessee, United States of America

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

William Richardson

Address Current address: Medical Research Council, Human Genetics Unit, Edinburgh, United Kingdom

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Georgina L Hamilton

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Robin C Allshire

To whom correspondence should be addressed. E-mail: Robin.Allshire@ed.ac.uk

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Alison L Pidoux

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

AGC, BGM, JFP, RCA, and ALP conceived and designed the experiments. AGC, BGM, JFP, WR, GLH, and ALP performed the experiments and contributed reagents/materials/analysis tools. AGC, BGM, JFP, WR, RCA, and ALP analyzed the data. AGC, RCA, and ALP wrote the paper.