About the Authors

Helen Liu

Affiliation Program in Epithelial Biology, Stanford University School of Medicine, Stanford, California, United States of America

Adam S Adler

Affiliation Program in Epithelial Biology, Stanford University School of Medicine, Stanford, California, United States of America

Eran Segal

Affiliation Department of Computer Science and Applied Mathematics, The Weizmann Institute of Science, Rehovot, Israel

Howard Y Chang

To whom correspondence should be addressed. E-mail: howchang@stanford.edu

Affiliation Program in Epithelial Biology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HL and HYC conceived and designed the experiments, analyzed the data, and wrote the paper. ASA analyzed the data and ES contributed analysis tools.