About the Authors

Chao-Tsung Yang

Affiliation Department of Genetics, Washington University School of Medicine, Saint Louis, Missouri, United States

Anna E Hindes

Affiliation Department of Genetics, Washington University School of Medicine, Saint Louis, Missouri, United States

Keith A Hultman

Affiliation Department of Genetics, Washington University School of Medicine, Saint Louis, Missouri, United States

Stephen L Johnson

To whom correspondence should be addressed. E-mail: sjohnson@genetics.wustl.edu

Affiliation Department of Genetics, Washington University School of Medicine, Saint Louis, Missouri, United States

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

CTY and SLJ conceived and designed the experiments. CTY, AEH, and KAH performed the experiments. CTY and SLJ analyzed the data and wrote the paper.