About the Authors

Gene W Yeo

To whom correspondence should be addressed. E-mail: geneyeo@salk.edu

Affiliations Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, United States of America , Laboratory of Genetics, Salk Institute, La Jolla, California, United States of America

Eric L. Van Nostrand

Address ¤ Current address: Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Affiliations Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, United States of America , Laboratory of Genetics, Salk Institute, La Jolla, California, United States of America

Tiffany Y Liang

Affiliations Crick-Jacobs Center for Theoretical and Computational Biology, Salk Institute, La Jolla, California, United States of America , Laboratory of Genetics, Salk Institute, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

GWY conceived and designed the experiments. GWY, ELVN, and TYL performed the experiments. GWY and ELVN analyzed the data. GWY wrote the paper.