About the Authors

Martha Susiarjo

Affiliations Department of Genetics, Case Western Reserve University, Cleveland, Ohio, United States of America , School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington, United States of America

Terry J Hassold

Affiliation School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington, United States of America

Edward Freeman

Affiliations Department of Genetics, Case Western Reserve University, Cleveland, Ohio, United States of America , Department of Biology, St. John Fisher College, Rochester, New York, United States of America

Patricia A Hunt

To whom correspondence should be addressed. E-mail: pathunt@wsu.edu

Affiliation School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MS and PAH conceived and designed the experiments. MS performed the experiments. MS, TJH, and PAH analyzed the data. TJH and PAH contributed reagents/materials/analysis tools. EF assisted in some methods used in studies. MS, TJH, and PAH wrote the paper.