About the Authors

Keyan Zhao, María José Aranzana, Sung Kim, Chunlao Tang, Christopher Toomajian, Honggang Zheng, Magnus Nordborg
Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Clare Lister, Chikako Shindo, Caroline Dean
Department of Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Paul Marjoram
Department of Preventive Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Corresponding Author

magnus@usc.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

C. Dean, P. Marjoram, and M. Nordborg designed and directed the overall project. M. J. Aranzana, C. Tang, C. Toomajian, and H. Zheng generated the candidate locus genotypes. M. J. Aranzana, C. Lister, C. Shindo, C. Tang, and H. Zheng generated the phenotypes. C. Lister and C. Shindo carried out the linkage mapping studies. K. Zhao, M. J. Aranzana, S. Kim, C. Toomajian, P. Marjoram, and M. Nordborg analyzed the data. K. Zhao and M. Nordborg wrote the paper.