About the Authors

Keyan Zhao

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

María José Aranzana

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Sung Kim

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Clare Lister

Affiliation Department of Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Chikako Shindo

Affiliation Department of Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Chunlao Tang

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Christopher Toomajian

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Honggang Zheng

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Caroline Dean

Affiliation Department of Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Paul Marjoram

Affiliation Department of Preventive Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Magnus Nordborg

To whom correspondence should be addressed. E-mail: magnus@usc.edu

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

C. Dean, P. Marjoram, and M. Nordborg designed and directed the overall project. M. J. Aranzana, C. Tang, C. Toomajian, and H. Zheng generated the candidate locus genotypes. M. J. Aranzana, C. Lister, C. Shindo, C. Tang, and H. Zheng generated the phenotypes. C. Lister and C. Shindo carried out the linkage mapping studies. K. Zhao, M. J. Aranzana, S. Kim, C. Toomajian, P. Marjoram, and M. Nordborg analyzed the data. K. Zhao and M. Nordborg wrote the paper.