About the Authors

Wenbo Ma

Affiliation Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada

Frederick F. T Dong

Affiliation Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada

John Stavrinides

Affiliation Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada

David S Guttman

To whom correspondence should be addressed. E-mail: david.guttman@utoronto.ca

Affiliations Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada, Centre for the Analysis of Genome Evolution and Function, University of Toronto, Toronto, Ontario, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

WM and DSG conceived and designed the experiments. WM and FFTD performed the experiments. JS and DSG analyzed the data. DSG contributed reagents/materials/analysis tools. WM, JS, and DSG wrote the paper.