About the Authors

Daniel J Gaffney

To whom correspondence should be addressed. E-mail: Daniel.Gaffney@mcgill.ca

Affiliations Institute of Evolutionary Biology, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom , McGill University and Genome Quebec Innovation Centre, Montréal, Québec, Canada

Peter D Keightley

Affiliation Institute of Evolutionary Biology, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DJG conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, and wrote the paper. PDK provided advice and comments on the manuscript.