About the Authors

Su-Chiung Fang

Affiliation Plant Biology Laboratory, The Salk Institute, La Jolla, California, United States of America

Chris de los Reyes

Affiliation Plant Biology Laboratory, The Salk Institute, La Jolla, California, United States of America

James G Umen

To whom correspondence should be addressed. E-mail: umen@salk.edu

Affiliation Plant Biology Laboratory, The Salk Institute, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SCF and JGU conceived and designed the experiments. SCF, CdlR, and JGU performed the experiments. SCF and JGU analyzed the data. SCF and JGU wrote the paper.