About the Authors

Haipeng Li

To whom correspondence should be addressed. E-mail: li@zi.biologie.uni-muenchen.de

Affiliation Section of Evolutionary Biology, Department of Biology II, University of Munich, Planegg-Martinsried, Germany

Wolfgang Stephan

Affiliation Section of Evolutionary Biology, Department of Biology II, University of Munich, Planegg-Martinsried, Germany

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HL and WS conceived the study and wrote the paper. HL designed and performed the analysis and simulations.