About the Authors

Michael F Seldin

To whom correspondence should be addressed. E-mail: mfseldin@ucdavis.edu

Affiliation Rowe Program in Human Genetics, Departments of Biological Chemistry and Medicine, University of California Davis, Davis, California, United States of America

Russell Shigeta

Affiliation Rowe Program in Human Genetics, Departments of Biological Chemistry and Medicine, University of California Davis, Davis, California, United States of America

Pablo Villoslada

Affiliation Center for Applied Medical Research, University of Navarra, Pamplona, Spain

Carlo Selmi

Affiliations Department of Medicine, Surgery, and Dentistry, San Paolo School of Medicine, University of Milan, Milan, Italy , Department of Medicine, University of California Davis, Davis, California, United States of America

Jaakko Tuomilehto

Affiliations Department of Epidemiology and Health Promotion, National Public Health Institute, Helsinki, Finland , South Ostrobothnia Central Hospital, Seinajoki, Finland

Gabriel Silva

Affiliation Obras Sociales del Hermano Pedro, Antigua, Guatemala

John W Belmont

Affiliation Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America

Lars Klareskog

Affiliation Karolinska University Hospital, Stockholm, Sweden

Peter K Gregersen

Affiliation The Robert S. Boas Center for Genomics and Human Genetics, Feinstein Institute for Medical Research, North Shore Long Island Jewish Health System, Manhasset, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The study was conceived by MFS and designed by MFS and PKG. MFS, RS, PV, CS, JT, GS, JWB, LK, and PKG recruited participants, and obtained and prepared DNA samples used in these studies. Analyses were performed by MFS and PKG. The manuscript was written by MFS and PKG with contributions from PV, CS, JT, GS, JWB, and LK.