About the Authors

Michael A Eberle

To whom correspondence should be addressed. E-mail: eberle@u.washington.edu

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Mark J Rieder

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Leonid Kruglyak

Affiliations Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America , Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Deborah A Nickerson

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MAE and MJR conceived and designed the experiments. MAE performed the experiments. MAE analyzed the data. MAE, MJR, LK, and DAN wrote the paper.