About the Authors

Simon W.-L Chan

Address Current address: Section of Plant Biology, University of California Davis, Davis, California, United States of America

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Ian R Henderson

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Xiaoyu Zhang

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Govind Shah

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Jason S.-C Chien

Address Current address: Department of Molecular and Cellular Biology, University of California Berkeley, Berkeley, California, United States of America

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Steven E Jacobsen

To whom correspondence should be addressed. E-mail: jacobsen@ucla.edu

Affiliations Department of Molecular, Cell, and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America , Howard Hughes Medical Institute, University of California Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SW-LC and SEJ conceived and designed the experiments. SW-LC, IRH, XZ, GS, JS-CC, and SEJ performed the experiments. SW-LC and SEJ analyzed the data. SW-LC, IRH, XZ, and SEJ contributed reagents/materials/analysis tools. SW-LC and SEJ wrote the paper.