About the Authors

Dan L Nicolae

To whom correspondence should be addressed. E-mail: nicolae@galton.uchicago.edu

Affiliation Department of Statistics, The University of Chicago, Chicago, Illinois, United States of America

Xiaoquan Wen

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Benjamin F Voight

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Nancy J Cox

Affiliations Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America , Department of Medicine, The University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DLN and NJC conceived and designed the experiments. XW performed the experiments. DLN analyzed the data. DLN, XW, and BFV contributed reagents/materials/analysis tools. DLN and NJC wrote the paper.