About the Authors

Shigeto Seno

Affiliation Department of Bioinformatic Engineering, Graduate School of Information Science and Technology, Osaka University, Osaka, Japan

Yoichi Takenaka

Affiliation Department of Bioinformatic Engineering, Graduate School of Information Science and Technology, Osaka University, Osaka, Japan

Chikatoshi Kai

Affiliation Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan

Jun Kawai

Affiliations Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan , Genome Science Laboratory, Discovery Research Institute, RIKEN Wako Institute, Wako, Japan

Piero Carninci

Affiliations Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan , Genome Science Laboratory, Discovery Research Institute, RIKEN Wako Institute, Wako, Japan

Yoshihide Hayashizaki

Affiliations Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan , Genome Science Laboratory, Discovery Research Institute, RIKEN Wako Institute, Wako, Japan

Hideo Matsuda

To whom correspondence should be addressed. E-mail: matsuda@ist.osaka-u.ac.jp

Affiliation Department of Bioinformatic Engineering, Graduate School of Information Science and Technology, Osaka University, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SS, YT, and HM conceived and designed the experiments. SS performed the experiments and analyzed the data. SS, CK, JK, PC, and YH contributed reagents/materials/analysis tools. SS, YT, and HM wrote the paper.