About the Authors

Masaaki Furuno

Contributed equally to this work with: Masaaki Furuno, Ken C Pang

Affiliation Mouse Genome Informatics Consortium, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Ken C Pang

Contributed equally to this work with: Masaaki Furuno, Ken C Pang

Affiliations Australian Research Council Special Research Centre for Functional and Applied Genomics, Institute for Molecular Bioscience, University of Queensland, Brisbane, Australia , T Cell laboratory, Ludwig Institute for Cancer Research, Austin Health, Heidelberg, Victoria, Australia

Noriko Ninomiya

Affiliation Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan

Shiro Fukuda

Affiliation Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan

Martin C Frith

Affiliations Australian Research Council Special Research Centre for Functional and Applied Genomics, Institute for Molecular Bioscience, University of Queensland, Brisbane, Australia , Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan

Carol Bult

Affiliation Mouse Genome Informatics Consortium, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Chikatoshi Kai

Affiliation Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan

Jun Kawai

Affiliations Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan , Genome Science Laboratory, Discovery Research Institute, RIKEN Wako Institute, Wako, Japan

Piero Carninci

Affiliations Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan , Genome Science Laboratory, Discovery Research Institute, RIKEN Wako Institute, Wako, Japan

Yoshihide Hayashizaki

Affiliations Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan , Genome Science Laboratory, Discovery Research Institute, RIKEN Wako Institute, Wako, Japan

John S Mattick

Affiliation Australian Research Council Special Research Centre for Functional and Applied Genomics, Institute for Molecular Bioscience, University of Queensland, Brisbane, Australia

Harukazu Suzuki

To whom correspondence should be addressed. E-mail: rgscerg@gsc.riken.jp

Affiliation Genome Exploration Research Group (Genome Network Project Core Group), RIKEN Genomic Sciences Center, RIKEN Yokohama Institute, Yokohama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MF, KCP, PC, YH, and HS conceived and designed the experiments. NN, SF, CK, and HS performed the experiments. MF, KCP, MCF, and HS analyzed the data. MF, CK, JK, PC, YH, and HS contributed reagents/materials/analysis tools. MF, KCP, CB, JSM, and HS wrote the paper.