About the Authors

Bettina Meier

Current address: School of Life Sciences, University of Dundee, Dundee, United Kingdom

Affiliations Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America , Wellcome Trust Biocenter, University of Dundee, Dundee, United Kingdom

Iuval Clejan

Affiliations Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America , Lineberger Comprehensive Cancer Center, University of North Carolina, Chapel Hill, North Carolina, United States of America

Yan Liu

Current address: Laboratory of Biochemistry and Genetics, National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Mia Lowden

Affiliations Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America , Department of Biology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Anton Gartner

Affiliation Wellcome Trust Biocenter, University of Dundee, Dundee, United Kingdom

Jonathan Hodgkin

Affiliation Genetics Unit, Department of Biochemistry, University of Oxford, Oxford, United Kingdom

Shawn Ahmed

To whom correspondence should be addressed. E-mail: shawn@med.unc.edu

Affiliations Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America , Lineberger Comprehensive Cancer Center, University of North Carolina, Chapel Hill, North Carolina, United States of America , Department of Biology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

BM, IC, ML, JH, and SA conceived and designed the experiments. BM, IC, YL, ML, and SA performed the experiments and analyzed the data. AG provided materials, lab space, and help with experiments. BM, IC, ML, JH, and SA wrote the paper.