About the Authors

Dennis C Ko

Affiliation Departments of Developmental Biology, Genetics, and Bioengineering, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Ljiljana Milenkovic

Affiliation Departments of Developmental Biology, Genetics, and Bioengineering, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Steven M Beier

Affiliation Departments of Developmental Biology, Genetics, and Bioengineering, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Hermogenes Manuel

Affiliation Departments of Developmental Biology, Genetics, and Bioengineering, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

JoAnn Buchanan

Affiliation Departments of Developmental Biology, Genetics, and Bioengineering, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Matthew P Scott

To whom correspondence should be addressed. E-mail: mscott@stanford.edu

Affiliation Departments of Developmental Biology, Genetics, and Bioengineering, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DCK, LM, SMB, and MPS conceived and designed the experiments. DCK, LM, HM, and JB performed the experiments. DCK analyzed the data. DCK, LM, SMB, HM, and JB contributed reagents/materials/analysis tools. DCK, LM, and MPS wrote the paper.