About the Authors

Bingshan Li

bingshan.li@vanderbilt.edu (BL); goncalo@umich.edu (GA)

Affiliation Center for Human Genetics Research, Department of Physiology and Biophysics, Vanderbilt University, Nashville, Tennessee, United States of America

Wei Chen

Affiliation Department of Pediatrics, Children's Hospital of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Xiaowei Zhan

Affiliation Center for Statistical Genetics, Department of Biostatistics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Fabio Busonero

Affiliations Center for Statistical Genetics, Department of Biostatistics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America, Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche (CNR), Monserrato, Italy

Serena Sanna

Affiliation Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche (CNR), Monserrato, Italy

Carlo Sidore

Affiliation Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche (CNR), Monserrato, Italy

Francesco Cucca

Affiliation Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche (CNR), Monserrato, Italy

Hyun M. Kang

Affiliation Center for Statistical Genetics, Department of Biostatistics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Gonçalo R. Abecasis

bingshan.li@vanderbilt.edu (BL); goncalo@umich.edu (GA)

Affiliation Center for Statistical Genetics, Department of Biostatistics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BL GRA. Performed the experiments: BL WC XZ. Analyzed the data: BL WC XZ CS. Contributed reagents/materials/analysis tools: FB SS CS FC HMK. Wrote the paper: BL GRA.