About the Authors

Lee B. Smith

Lee.Smith@ed.ac.uk

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Laura Milne

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Nancy Nelson

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Sharon Eddie

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Pamela Brown

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Nina Atanassova

Affiliation Institute of Experimental Morphology and Anthropology with Museum, Bulgarian Academy of Sciences, Sofia, Bulgaria

Moira K. O'Bryan

Affiliation The Department of Anatomy and Developmental Biology, Monash University, Melbourne, Victoria, Australia

Liza O'Donnell

Affiliations The Department of Anatomy and Developmental Biology, Monash University, Melbourne, Victoria, Australia, Prince Henry's Institute of Medical Research, Clayton, Victoria, Australia

Danielle Rhodes

Affiliation The Department of Anatomy and Developmental Biology, Monash University, Melbourne, Victoria, Australia

Sara Wells

Affiliation Mary Lyon Centre, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom

Diane Napper

Affiliation Mary Lyon Centre, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom

Patrick Nolan

Affiliation Mammalian Genetics Unit, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom

Zuzanna Lalanne

Affiliation Mammalian Genetics Unit, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom

Michael Cheeseman

Affiliations Mary Lyon Centre, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom, Mammalian Genetics Unit, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom

Josephine Peters

Affiliation Mammalian Genetics Unit, Medical Research Council, Harwell Science and Innovation Campus, Harwell, Oxfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LBS MKO JP. Performed the experiments: LBS LM NN SE PB NA DR SW DN ZL MC. Analyzed the data: LBS LM MKO LO JP. Contributed reagents/materials/analysis tools: PN. Wrote the paper: LBS.