About the Authors

Oliver P. Forman

oliver.forman@aht.org.uk

Affiliation Kennel Club Genetics Centre, Animal Health Trust, Kentford, United Kingdom

Jacques Penderis

Contributed equally to this work with: Jacques Penderis, Claudia Hartley

Affiliation The School of Veterinary Medicine, University of Glasgow, Glasgow, United Kingdom

Claudia Hartley

Contributed equally to this work with: Jacques Penderis, Claudia Hartley

Affiliation Kennel Club Genetics Centre, Animal Health Trust, Kentford, United Kingdom

Louisa J. Hayward

Affiliation Kennel Club Genetics Centre, Animal Health Trust, Kentford, United Kingdom

Sally L. Ricketts

Affiliation Kennel Club Genetics Centre, Animal Health Trust, Kentford, United Kingdom

Cathryn S. Mellersh

Affiliation Kennel Club Genetics Centre, Animal Health Trust, Kentford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: OPF JP CH LJH SLR CSM. Performed the experiments: OPF LJH. Analyzed the data: OPF LJH SLR. Contributed reagents/materials/analysis tools: OPF JP CH SLR LJH. Wrote the paper: OPF JP CH SLR LJH CSM.