About the Authors

Zhenming Yu

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Xiuyin Teng

Affiliations Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Howard Hughes Medical Institute, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Nancy M. Bonini

nbonini@sas.upenn.edu

Affiliations Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Howard Hughes Medical Institute, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZY NMB. Performed the experiments: ZY XT. Analyzed the data: ZY NMB. Wrote the paper: ZY NMB.