About the Authors

Yoshimi Tokuzawa

Contributed equally to this work with: Yoshimi Tokuzawa, Ken Yagi

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Ken Yagi

Contributed equally to this work with: Yoshimi Tokuzawa, Ken Yagi

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Yzumi Yamashita

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Yutaka Nakachi

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Itoshi Nikaido

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Hidemasa Bono

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Yuichi Ninomiya

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Yukiko Kanesaki-Yatsuka

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Masumi Akita

Affiliation Division of Morphological Science, Biomedical Research Center, Saitama Medical University, Iruma-gun, Saitama, Japan

Hiromi Motegi

Affiliation RIKEN BioResource Center, Tsukuba, Ibaraki, Japan

Shigeharu Wakana

Affiliation RIKEN BioResource Center, Tsukuba, Ibaraki, Japan

Tetsuo Noda

Affiliations RIKEN BioResource Center, Tsukuba, Ibaraki, Japan, The Cancer Institute of the Japanese Foundation for Cancer Research, Koto-ward, Tokyo, Japan

Fred Sablitzky

Affiliation Developmental Genetics and Gene Control, Institute of Genetics, University of Nottingham, Queen's Medical Center, Nottingham, United Kingdom

Shigeki Arai

Affiliation Division of Gene Structure and Function, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Riki Kurokawa

Affiliation Division of Gene Structure and Function, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Toru Fukuda

Affiliation Division of Pathophysiology, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Takenobu Katagiri

Affiliation Division of Pathophysiology, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Christian Schönbach

Affiliations Division of Genomics and Genetics, Nanyang Technological University School of Biological Sciences, Singapore, Singapore, Department of Bioscience and Bioinformatics, Kyushu Institute of Technology, Iizuka, Fukuoka, Japan

Tatsuo Suda

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Yosuke Mizuno

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Yasushi Okazaki

okazaki@saitama-med.ac.jp

Affiliation Division of Functional Genomics and Systems Medicine, Research Center for Genomic Medicine, Saitama Medical University, Hidaka, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Y Tokuzawa, K Yagi, Y Mizuno, Y Okazaki. Performed the experiments: Y Tokuzawa, K Yagi, Y Yamashita, Y Ninomiya, Y Kanesaki-Yatsuka, M Akita, H Motegi, S Wakana, T Noda, Y Mizuno. Analyzed the data: Y Tokuzawa, Y Yamashita, Y Nakachi, I Nikaido, H Bono. Contributed reagents/materials/analysis tools: F Sablitzky, S Arai, R Kurokawa, T Fukuda, T Katagiri. Wrote the paper: Y Tokuzawa, K Yagi, Y Nakachi, C Schönbach, T Suda, Y Okazaki.