About the Authors

Heather E. Wheeler

Affiliation Department of Genetics, Stanford University Medical Center, Stanford, California, United States of America

E. Jeffrey Metter

Affiliations Longitudinal Studies Section, Clinical Research Branch, National Institute on Aging, Baltimore, Maryland, United States of America, Medstar Research Institute, Baltimore, Maryland, United States of America

Toshiko Tanaka

Affiliations Longitudinal Studies Section, Clinical Research Branch, National Institute on Aging, Baltimore, Maryland, United States of America, Medstar Research Institute, Baltimore, Maryland, United States of America

Devin Absher

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

John Higgins

Affiliation Department of Pathology, Stanford University Medical Center, Stanford, California, United States of America

Jacob M. Zahn

Affiliation Stanford Genome Technology Center, Palo Alto, California, United States of America

Julie Wilhelmy

Affiliation Stanford Genome Technology Center, Palo Alto, California, United States of America

Ronald W. Davis

Affiliation Stanford Genome Technology Center, Palo Alto, California, United States of America

Andrew Singleton

Affiliation Laboratory of Neurogenetics, National Institute on Aging, Bethesda, Maryland, United States of America

Richard M. Myers

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Luigi Ferrucci

Affiliations Longitudinal Studies Section, Clinical Research Branch, National Institute on Aging, Baltimore, Maryland, United States of America, Medstar Research Institute, Baltimore, Maryland, United States of America

Stuart K. Kim

kim@cmgm.stanford.edu

Affiliations Department of Genetics, Stanford University Medical Center, Stanford, California, United States of America, Department of Developmental Biology, Stanford University Medical Center, Stanford, California, United States of America

Competing Interests

Two of the authors (HEW and SKK) are listed as the inventors in a patent filed by the Stanford University Office of Technology Licensing titled: A Genetic Test for Healthy Kidney Aging (Docket 08-360).

Author Contributions

Conceived and designed the experiments: HEW SKK. Performed the experiments: HEW JW. Analyzed the data: HEW EJM JMZ SKK. Contributed reagents/materials/analysis tools: EJM TT DA JH RWD AS RMM LF SKK. Wrote the paper: HEW SKK.