About the Authors

Chia Huey Ooi

Affiliation Duke-NUS Graduate Medical School, Singapore

Tatiana Ivanova

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Jeanie Wu

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Minghui Lee

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Iain Beehuat Tan

Affiliation Division of Medical Oncology, National Cancer Centre, Singapore

Jiong Tao

Affiliations Cellular and Molecular Research, National Cancer Centre, Singapore, Department of Physiology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore

Lindsay Ward

Affiliation Section of Pathology and Tumour Biology, Leeds Institute of Molecular Medicine, St. James's University Hospital, Leeds, United Kingdom

Jun Hao Koo

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Veena Gopalakrishnan

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Yansong Zhu

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Lai Ling Cheng

Affiliation Singapore-MIT Alliance, National University of Singapore, Singapore

Julian Lee

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Sun Young Rha

Affiliation Department of Internal Medicine, Yonsei Cancer Center, Yonsei University College of Medicine, Seoul, Korea

Hyun Cheol Chung

Affiliation Department of Internal Medicine, Yonsei Cancer Center, Yonsei University College of Medicine, Seoul, Korea

Kumaresan Ganesan

Affiliation Cellular and Molecular Research, National Cancer Centre, Singapore

Jimmy So

Affiliation Department of Surgery, Yong Loo Lin School of Medicine, National University of Singapore, Singapore

Khee Chee Soo

Affiliation Division of Surgical Oncology, National Cancer Centre, Singapore

Dennis Lim

Affiliation Department of General Surgery, Singapore General Hospital, Singapore

Weng Hoong Chan

Affiliation Department of General Surgery, Singapore General Hospital, Singapore

Wai Keong Wong

Affiliation Department of General Surgery, Singapore General Hospital, Singapore

David Bowtell

Affiliation Cancer Genomics and Biochemistry Laboratory, Peter MacCallum Cancer Centre, East Melbourne, Victoria, Australia

Khay Guan Yeoh

Affiliation Department of Medicine, Yong Loo Lin School of Medicine, National University of Singapore, Singapore

Heike Grabsch

Affiliation Section of Pathology and Tumour Biology, Leeds Institute of Molecular Medicine, St. James's University Hospital, Leeds, United Kingdom

Alex Boussioutas

Affiliations Cancer Genomics and Biochemistry Laboratory, Peter MacCallum Cancer Centre, East Melbourne, Victoria, Australia, Department of Medicine (RMH/WH), University of Melbourne, Western Hospital, Footscray, Victoria, Australia

Patrick Tan

gmstanp@duke-nus.edu.sg

Affiliations Duke-NUS Graduate Medical School, Singapore, Cellular and Molecular Research, National Cancer Centre, Singapore, Cancer Science Institute of Singapore, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Genome Institute of Singapore, Singapore

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CHO PT. Performed the experiments: CHO TI JW ML JT LW LLC KG HG. Analyzed the data: CHO TI HG. Contributed reagents/materials/analysis tools: TI JW ML IBT JHK VG YZ JL SYR HCC KG JS KCS DL WHC WKW DB KGY HG AB. Wrote the paper: CHO PT.