About the Authors

David Reich

reich@genetics.med.harvard.edu (DR); James.Wilson3@va.gov (JGW)

Affiliations Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America, Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Michael A. Nalls

Affiliations Laboratory of Neurogenetics, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America, Laboratory of Epidemiology, Demography and Biometry, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America

W. H. Linda Kao

Affiliation Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Ermeg L. Akylbekova

Affiliation Jackson Heart Study Analysis Group, Jackson State University, Jackson, Mississippi, United States of America

Arti Tandon

Affiliations Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America, Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Nick Patterson

Affiliation Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

James Mullikin

Affiliation Comparative Genomics Unit, Genome Technology Branch, National Human Genome Research Institute, Rockville, Maryland, United States of America

Wen-Chi Hsueh

Affiliation Division of Medical Genetics, Department of Medicine, Department of Epidemiology and Biostatistics, Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Ching-Yu Cheng

Affiliations Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America, Inherited Disease Research Branch, National Human Genome Research Institute, Baltimore, Maryland, United States of America

Josef Coresh

Affiliation Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Eric Boerwinkle

Affiliation Human Genetics Center, University of Texas Health Science Center at Houston, Houston, Texas, United States of America

Man Li

Affiliation Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Alicja Waliszewska

Affiliations Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America, Laboratory of Molecular Immunology, Center for Neurologic Disease, Brigham and Women's Hospital, Boston, Massachusetts, United States of America

Julie Neubauer

Affiliation Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Rongling Li

Affiliation Department of Preventive Medicine, Center for Genomics and Bioinformatics, University of Tennessee Health Science Center, Memphis, Tennessee, United States of America

Tennille S. Leak

Affiliation Department of Epidemiology, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Lynette Ekunwe

Affiliation Jackson Heart Study Analysis Group, Jackson State University, Jackson, Mississippi, United States of America

Joe C. Files

Affiliation Department of Medicine, Division of Hematology, University of Mississippi Medical Center, Jackson, Mississippi, United States of America

Cheryl L. Hardy

Affiliation Department of Medicine, Division of Hematology, University of Mississippi Medical Center, Jackson, Mississippi, United States of America

Joseph M. Zmuda

Affiliation Department of Epidemiology, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Herman A. Taylor

Affiliations Jackson State University, Jackson, Mississippi, United States of America, Tougaloo College, Jackson, Mississippi, United States of America, University of Mississippi Medical Center, Jackson, Mississippi, United States of America

Elad Ziv

Affiliations Division of General Internal Medicine, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America, Department of Epidemiology and Biostatistics, Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America, Helen Diller Family Comprehensive Cancer Center, University of California San Francisco, San Francisco, California, United States of America

Tamara B. Harris

Affiliation Laboratory of Epidemiology, Demography and Biometry, Intramural Research Program, National Institute on Aging, Bethesda, Maryland, United States of America

James G. Wilson

reich@genetics.med.harvard.edu (DR); James.Wilson3@va.gov (JGW)

Affiliations V.A. Medical Center, Jackson, Mississippi, United States of America, University of Mississippi Medical Center, Jackson, Mississippi, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DR EZ TBH JGW. Performed the experiments: DR MAN ELA AT AW JN JGW. Analyzed the data: DR MAN WHLK ELA AT NP JM ML JGW. Contributed reagents/materials/analysis tools: DR MAN WHLK ELA AT NP JM WCH CYC JC EB RL TSL LE JCF CLH JMZ HAT EZ TBH JGW. Wrote the paper: DR JGW.