About the Authors

J. Kim Holloway

Affiliation Department of Biomedical Sciences, Cornell University, Ithaca, New York, United States of America

James Booth

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Winfried Edelmann

Affiliation Department of Cell Biology, Albert Einstein College of Medicine, Bronx, New York, United States of America

Clare H. McGowan

Affiliation The Scripps Research Institute, La Jolla, California, United States of America

Paula E. Cohen

paula.cohen@cornell.edu

Affiliation Department of Biomedical Sciences, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JKH PEC. Performed the experiments: JKH. Analyzed the data: JKH JB PEC. Contributed reagents/materials/analysis tools: WE CHM PEC. Wrote the paper: JKH PEC.