About the Authors

Carolin Kosiol

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Tomáš Vinař

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Rute R. da Fonseca

Affiliation Institute of Biology, University of Copenhagen, Copenhagen, Denmark

Melissa J. Hubisz

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Carlos D. Bustamante

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Rasmus Nielsen

Affiliation Institute of Biology, University of Copenhagen, Copenhagen, Denmark

Adam Siepel

acs4@cornell.edu

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CK TV CDB RN AS. Performed the experiments: CK TV AS. Analyzed the data: CK TV RRdF RN AS. Contributed reagents/materials/analysis tools: CK TV MJH RN AS. Wrote the paper: CK TV AS.