About the Authors

Pierre Fontanillas

Affiliation Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Daniel L Hartl

Affiliation Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Max Reuter

To whom correspondence should be addressed. E-mail: m.reuter@ucl.ac.uk

Affiliation The Galton Laboratory, Department of Biology, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PF and MR conceived and designed the experiments. PF and MR analyzed the data. DLH contributed reagents/materials/analysis tools. PF, DLH, and MR wrote the paper.