About the Authors

Catharine E Boothroyd

Contributed equally to this work with: Catharine E Boothroyd, Herman Wijnen

Affiliation Laboratory of Genetics, The Rockefeller University, New York, New York, United States of America

Herman Wijnen

Contributed equally to this work with: Catharine E Boothroyd, Herman Wijnen

Affiliations Laboratory of Genetics, The Rockefeller University, New York, New York, United States of America , Department of Biology, University of Virginia, Charlottesville, Virginia, United States of America

Felix Naef

Affiliations Laboratory of Mathematical Physics, The Rockefeller University, New York, New York, United States of America , School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne, Lausanne, Switzerland

Lino Saez

Affiliation Laboratory of Genetics, The Rockefeller University, New York, New York, United States of America

Michael W Young

To whom correspondence should be addressed. E-mail: young@mail.rockefeller.edu

Affiliation Laboratory of Genetics, The Rockefeller University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

All authors conceived and designed the experiments and analyzed the data. CEB and HW performed the experiments. CEB, HW, and FN contributed reagents/materials/analysis tools. CEB, HW, FN, and MWY wrote the paper.